Chapter 53

1) A population is correctly defined as having which of the following characteristics?
I. inhabiting the same general area
II. individuals belonging to the same species
III. possessing a constant and uniform density and dispersion
A) I only
B) III only
C) I and II only
D) II and III only
E) I, II, and III
2) An ecologist recorded 12 white-tailed deer, Odocoileus virginianus, per square mile in one
woodlot and 20 per square mile on another woodlot. What was the ecologist comparing?
A) density
B) dispersion
C) carrying capacity
D) quadrats
E) range
3) To measure the population density of monarch butterflies occupying a particular park, 100
butterflies are captured, marked with a small dot on a wing, and then released. The next
day, another 100 butterflies are captured, including the recapture of 20 marked butterflies.
One would estimate the population to be
A) 200.
B) 500.
C) 1,000.
D) 10,000.
E) 900,000.
4) During the spring, you are studying the mice that live in a field near your home. There are
lots of mice in this field, but you realize that you rarely observe any reproductive females.
This most likely indicates
A) that there is selective predation on female mice.
B) that female mice die before reproducing.
C) that this habitat is a good place for mice to reproduce.
D) that you are observing immigrant mice.
E) that the breeding season is over
5) You are observing a population of lizards when you notice that the number of adults has
increased and is higher than previously observed. One explanation for such an observation
would include
A) reduction in death rate.
B) increased immigration.
C) increased emigration.
D) decreased emigration.
E) increased birth rate.
6) The most common kind of dispersion in nature is
A) clumped.
B) random.
C) uniform.
D) indeterminate.
E) dispersive.
7) Uniform spacing patterns in plants such as the creosote bush are most often associated with
A) chance.
B) patterns of high humidity.
C) the random distribution of seeds.
D) competitive interactions among individuals in the population.
E) the concentration of nutrients within the populationʹs range.
8) Which of the following groups would be most likely to exhibit uniform dispersion?
A) red squirrels, who actively defend territories
B) cattails, which grow primarily at edges of lakes and streams
C) dwarf mistletoes, which parasitize particular species of forest tree
D) moths in a city at night
E) lake trout, which seek out deep water
9) A table listing such items as age, observed number of organisms alive each year, and life
expectancy is known as a (an)
A) life table.
B) mortality table.
C) survivorship table.
D) rate table.
E) insurance table.
10) Life tables are most useful in determining which of the following?
A) carrying capacity
B) the fate of a cohort of newborn organisms throughout their lives
C) immigration and emigration rates
D) population dispersion patterns
E) reproductive rates
17) In order to construct a reproductive table for a sexual species, you need to
A) assess sperm viability.
B) keep track of all of the offspring of a cohort.
C) keep track of the females in a cohort.
D) keep track of all of the offspring of the females in a cohort.
E) keep track of the ratio of deaths to births in a cohort.
18) Which of the following examples would most accurately measure the density of the
population being studied?
A) counting the number of prairie dog burrows per hectare
B) counting the number of times a 1 kilometer transect is intersected by tracks of red
squirrels after a snowfall
C) counting the number of coyote droppings per hectare
D) multiplying the number of moss plants counted in 10, 1m2 quadrats by 100 to
determine the density per kilometer2.
E) counting the number of zebras from airplane census observations.
19) To measure the population of lake trout in a 250 hectare lake, 200 individuals were netted
and marked with a fin clip, and then returned to the lake. The next week, the lake is netted
again, and out of the 200 lake trout that are caught, 50 have fin clips. Using the
capture-recapture estimate, the lake trout population size could be closest to which of the
A) 200
B) 250
C) 400
D) 800
E) 40,000
20) Which of the following assumptions have to be made regarding the capture -recapture
estimate of population size?
I. Marked and unmarked individuals have the same probability of being trapped.
II. The marked individuals have thoroughly mixed with population after being
III. No individuals have entered or left the population by immigration or emigration,
and no individuals have been added by birth or eliminated by death during the course of
the estimate.
A) I only
B) II only
C) I and II only
D) II and III only
E) I, II, and III
21) Long-term studies of Beldingʹs ground squirrels show that immigrants move nearly 2 km
from where they are born and make up 1 to 8% of the males and 0.7 to 6% of the females in
other populations. On an evolutionary scale, why is this significant?
A) These immigrants make up for the deaths of individuals keeping the other
populationsʹ size stable.
B) Young reproductive males tend to stay in their home population and are not driven
out by other territorial males.
C) These immigrants provide a source of genetic diversity for the other populations.
D) Those individuals that emigrate to these new populations are looking for less
crowded conditions with more resources.
E) Gradually, the populations of ground squirrels will move from a uniform to a
clumped population pattern of dispersion.
22) Demography is the study of
A) the vital statistics of populations and how they change over time.
B) death and emigration rates of a population at any moment in time.
C) the survival patterns of a population.
D) life expectancy of individuals within a population.
E) reproductive rates of a population during a given year.
23) Natural selection has led to the evolution of diverse natural history strategies, which have
in common
A) many offspring per reproductive episode.
B) limitation only by density-independent limiting factors.
C) adaptation to stable environments.
D) maximum lifetime reproductive success.
E) relatively large offspring.
24) Natural selection involves energetic trade-offs between
A) choosing how many offspring to produce over the course of a lifetime and how long
to live.
B) producing large numbers of gametes when employing internal fertilization versus
fewer numbers of gametes when employing external fertilization.
C) the emigration of individuals when they are no longer reproductively capable or
committing suicide.
D) increasing the number of individuals produced during each reproductive episode
with a corresponding decrease in parental care.
E) high survival rates of offspring and the cost of parental care.
25) The three basic variables that make up the life history of an organism are
A) life expectancy, birth rate, and death rate.
B) number of reproductive females in the population, age structure of the population,
and life expectancy.
C) age when reproduction begins, how often reproduction occurs, and how many
offspring are produced per reproductive episode.
D) how often reproduction occurs, life expectancy of females in the population, and
number of offspring per reproductive episode.
E) the number of reproductive females in the population, how often reproduction
occurs, and death rate.
Researchers in the Netherlands studied the effects of parental caregiving in European kestrels over 5 years. The researchers transferred chicks among nests to produce reduced broods (three or four chicks),
normal broods (five or six), and enlarged broods (seven or eight). They then measured the percentage of
male and female parent birds that survived the following winter. (Both males and females provide care
for chicks.)

26) Which of the following is a conclusion that can be drawn from this graph?
A) Female survivability is more negatively affected by larger brood size than is male
B) Male survivability decreased by 50% between reduced and enlarged brood
C) Both males and females had increases in daily hunting with the enlarged brood size.
D) There appears to be a negative correlation between brood enlargements and parental
E) Chicks in reduced brood treatment received more food, weight gain, and reduced

27) Which of the following pairs of reproductive strategies is consistent with energetic
trade-off and reproductive success?
A) Pioneer species of plants produce many very small, highly airborne seeds, while large
elephants that are very good parents produce many offspring.
B) Female rabbits that suffer high predation rates may produce several litters per
breeding season, and coconuts produce few fruits, but most survive when they
encounter proper growing conditions.
C) Species that have to broadcast to distant habitats tend to produce seeds with heavy
protective seed coats, and animals that are caring parents produce fewer offspring
with lower infant mortality.
D) Free-living insects lay thousands of eggs and provide no parental care, while flowers
take good care of their seeds until they are ready to germinate.
E) Some mammals will not reproduce when environmental resources are low so they
can survive until conditions get better, and plants that produce many small seeds are
likely found in stable environments.
28) A population of ground squirrels has an annual per capita birth rate of 0.06 and an annual
per capita death rate of 0.02. Estimate the number of individuals added to (or lost from) a
population of 1,000 individuals in one year.
A) 120 individuals added
B) 40 individuals added
C) 20 individuals added
D) 400 individuals added
E) 20 individuals lost
29) A small population of white-footed mice has the same intrinsic rate of increase (r) as a large
population. If everything else is equal,
A) the large population will add more individuals per unit time.
B) the small population will add more individuals per unit time.
C) the two populations will add equal numbers of individuals per unit time.
D) the J-shaped growth curves will look identical.
E) the growth trajectories of the two populations will proceed in opposite directions.
30) Imagine that you are managing a large ranch. You know from historical accounts that wild
sheep used to live there, but they have been extirpated. You decide to reintroduce them.
After doing some research to determine what might be an appropriately sized founding
population, you do so. You then watch the population increase for several generations, and
graph the number of individuals (vertical axis) against the number of generations
(horizontal axis). The graph will appear as
A) a diagonal line, getting higher with each generation.
B) an ʺS,ʺ increasing with each generation.
C) an upside-down ʺU.ʺ
D) a ʺJ,ʺ increasing with each generation.
E) an ʺSʺ that ends with a vertical line.
31) In the logistic equation dN/dt = rN (K-N)
, r is a measure of the populationʹs intrinsic rate of
increase. It is determined by which of the following?
A) birth rate and death rates
B) dispersion
C) density
D) carrying capacity
E) life history
32) In 2005, the United States had a population of approximately 295,000,000 people. If the
birth rate was 13 births for every 1,000 people, approximately how many births occurred in
the United States in 2005?
A) 3,800
B) 38,000
C) 380,000
D) 3,800,000
E) 38,000,000
35) As N approaches K for a certain population, which of the following is predicted by the
logistic equation?
A) The growth rate will not change.
B) The growth rate will approach zero.
C) The population will show an Allee effect.
D) The population will increase exponentially.
E) The carrying capacity of the environment will increase.
36) Often the growth cycle of one population has an effect on the cycle of another. As moose
populations increase, wolf populations also increase. Thus, if we are considering the logistic
equation for the wolf population,
dN/dt = rN (K-N)
which of the factors accounts for the effect on the moose population?
A) r
B) N
C) rN
D) K
E) dt
37) Which of the following might be expected in the logistic model of population growth?
A) As N approaches K, b increases.
B) As N approaches K, r increases.
C) As N approaches K, d increases.
D) Both A and B are true.
E) Both B and C are true.
38) In models of sigmoidal (logistic) population growth,
A) population growth rate slows dramatically as N approaches K.
B) new individuals are added to the population most rapidly at the beginning of the
populationʹs growth.
C) only density-dependent factors affect the rate of population growth.
D) only density-independent factors affect the rate of population growth.
E) carrying capacity is never reached.
39) The Allee effect is used to describe a population that
A) has become so small that it will have difficulty surviving and reproducing.
B) has become so large it will have difficulty surviving and reproducing.
C) approaches carrying capacity.
D) exceeds carrying capacity.
E) is in crash decline.
40) Which of the following is the pattern of spacing for individuals within the boundaries of
the population?
A) cohort
B) dispersion
C) Allee effect
D) iteroparous
E) semelparous
41) Pacific salmon or annual plants illustrate which of the following?
A) cohort
B) dispersion
C) Allee effect
D) iteroparous
E) semelparous
42) Which of the following describes having more than one reproductive episode during a
A) cohort
B) dispersion
C) Allee effect
D) iteroparous
E) semelparous
43) Density-dependent factors are related to which of the following?
A) cohort
B) dispersion
C) Allee effect
D) iteroparous
E) semelparous
44) Which of the following is true?
A) K-selection operates in populations where populations fluctuate well below the
carrying capacity.
B) r-selection occurs in populations whose densities are very near the carrying capacity.
C) Different populations of the same species will be consistently r- or K-selected.
D) r- and K-selection are two extremes of a range of life history strategies.
E) r-selection tends to maximize population size, not the rate of increase in population
45) The life history traits favored by selection are most likely to vary with
A) fluctuations in K.
B) the shape of the J curve.
C) the maximum size of a population.
D) population density.
E) population dispersion.
46) In which of the following habitats would you expect to find the largest number of
K-selected individuals?
A) a recently abandoned agricultural field in Ohio
B) the sand dune communities of south Lake Michigan
C) the flora and fauna of a coral reef in the Caribbean
D) South Florida after a hurricane
E) a newly emergent volcanic island
47) Which of the following characterizes relatively K-selected populations?
A) offspring with good chances of survival
B) many offspring per reproductive episode
C) small offspring
D) a high intrinsic rate of increase
E) early parental reproduction
48) Which of the following statements about the evolution of life histories is correct?
A) Stable environments with limited resources favor r-selected populations.
B) K-selected populations are most often found in environments where
density-independent factors are important regulators of population size.
C) Most populations have both r- and K-selected characteristics that vary under
different environmental conditions.
D) The reproductive efforts of r-selected populations are directed at producing just a few
offspring with good competitive abilities.
E) K-selected populations rarely approach carrying capacity.
49) Your friend comes to you with a problem. It seems his shrimp boats arenʹt catching nearly
as much shrimp as they used to. He canʹt understand why because originally he caught all
the shrimp he could handle. Each year he added a new boat, and for a long time each boat
caught tons of shrimp. As he added more boats, there came a time when each boat caught
somewhat fewer shrimp, and now, each boat is catching a lot less shrimp. Which of the
following topics might help your friend understand the source of his problem?
A) density-dependent population regulation and intrinsic characteristics of population
B) exponential growth curves and unlimited environmental resources
C) density-independent population regulation and chance occurrence
D) pollution effects of a natural environment and learned shrimp behavior
E) a K-selected population switching to an r-selected population
50) Carrying capacity is
A) seldom reached by marine producers and consumers because of the vast resources of
the ocean.
B) the maximum population size that a particular environment can support.
C) fixed for most species over most of their range most of the time.
D) determined by density and dispersion data.
E) the term used to describe the stress a population undergoes due to limited resources.
56) Which of the following can contribute to density-dependent regulation of populations?
A) the removal of toxic waste by decomposers
B) intraspecific competition for nutrients
C) earthquakes
D) floods
E) weather catastrophes
57) Field observation suggests that populations of a particular species of herbivorous mammal
undergo cyclic fluctuations in density at three- to five-year intervals. Which of the
following represent (a) plausible explanation(s) of these cycles?
A) Periodic crowding affects the endocrine system, resulting in increased aggressiveness.
B) Increases in population density lead to increased rates of predation.
C) Increases in rates of herbivory lead to changes in the nutritive value of plants used as
D) Increases in population density lead to more proximal infestations of parasites to host
E) All of the above are plausible explanations of population cycling.
58) Which of the following is an incorrect statement about the regulation of populations?
A) The logistic equation reflects the effect of density-dependent factors, which can
ultimately stabilize populations around the carrying capacity.
B) Density-independent factors have an increasingly greater effect as a populationʹs
density increases.
C) High densities in a population may cause physiological changes that inhibit
D) Because of the overlapping nature of population-regulating factors, it is often difficult
to precisely determine their cause-and-effect relationships.
E) The occurrence of population cycles in some populations may be the result of
crowding or lag times in the response to density-dependent factors.
59) Which of the following is a density-independent factor limiting human population growth?
A) social pressure for birth control
B) earthquakes
C) plagues
D) famines
E) pollution
60) A population of white-footed mice becomes severely overpopulated in a habitat that has
been disturbed by human activity. Sometimes intrinsic factors cause the population to
increase in mortality and lower reproduction rates in reaction to the stress of
overpopulation. Which of the following is an example of intrinsic population control?
A) Owl populations frequent the area more often because of increased hunting success.
B) Females undergo hormonal changes that delay sexual maturation and many
individuals suffer depressed immune systems and die due to the stress of
C) Clumped dispersion of the population leads to increased spread of disease and
parasites resulting in a population crash.
D) All of the resources (food and shelter) are used up by overpopulation and much of the
population dies of exposure and/or starvation.
E) Because the individuals are vulnerable they are more likely to die off if a drought or
flood were to occur.
62) Consider several human populations of equal size and net reproductive rate, but different
in age structure. The population that is likely to grow the most during the next 30 years is
the one with the greatest fraction of people in which age range?
A) 50 to 60 years
B) 40 to 50 years
C) 30 to 40 years
D) 20 to 30 years
E) 10 to 20 years
67) Most ecologists believe that the average global carrying capacity for the human population
is between
A) 5 and 6 billion.
B) 6 and 8 billion.
C) 10 and 15 billion.
D) 15 and 20 billion.
E) 20 and 25 billion.
68) An ecological footprint is a construct that is useful
A) for a person living in a developed nation to consider to make better choices when
using global food and energy resources.
B) for a person living in a developing country to see how much of the worldʹs resources
are left for him/her.
C) in converting human foodsʹ meat biomass to plant biomass.
D) in making predictions about the global carrying capacity of humans.
E) in determining which nations produce the least amount of carbon dioxide from the
burning of fossil fuels.
1) The observation that members of a population are uniformly distributed suggests that
A) the size of the area occupied by the population is increasing.
B) resources are distributed unevenly.
C) the members of the population are competing for access to a resource.
D) the members of the population are neither attracted to nor repelled by one another.
E) the density of the population is low.
2) Population ecologists follow the fate of same-age cohorts to
A) determine a populationʹs carrying capacity.
B) determine if a population is regulated by density-dependent processes.
C) determine the birth rate and death rate of each group in a population.
D) determine the factors that regulate the size of a population.
E) determine if a populationʹs growth is cyclic.
3) According to the logistic growth equation dN/dt = rmaxN(K — N)/K,
A) the number of individuals added per unit time is greatest when N is close to zero.
B) the per capita growth rate (r) increases as N approaches K.
C) population growth is zero when N equals K.
D) the population grows exponentially when K is small.
E) the birth rate (b) approaches zero as N approaches K.
4) A populationʹs carrying capacity
A) can be accurately calculated using the logistic growth model.
B) generally remains constant over time.
C) increases as the per capita growth rate (r) decreases.
D) may change as environmental conditions change.
E) can never be exceeded.
5) Which pair of terms most accurately describes life history traits for a stable population of
A) Semelparous; r-selected
B) Semelparous; K-selected
C) Iteroparous; r-selected
D) Iteroparous; K-selected
E) Iteroparous; N-selected
6) During exponential growth, a population always
A) grows by thousands of individuals.
B) grows at its maximum per capita rate.
C) quickly reaches its carrying capacity.
D) cycles through time.
E) loses some individuals to emigration.
7) Scientific study of the population cycles of the snowshoe hare and its predator, the lynx,
has revealed that
A) the prey population is controlled by predators alone.
B) hares and lynx are so mutually dependent that each species cannot survive without
the other.
C) multiple biotic and abiotic factors contribute to the cycling of hare and lynx
D) both hare and lynx populations are regulated mainly by abiotic factors.
E) the hare population is r-selected and the lynx population is K-selected.
8) Based on current growth rates, Earthʹs human population in 2010 will be closest to
A) 2 million.
B) 3 billion.
C) 4 billion.
D) 7 billion.
E) 10 billion.
9) Which of the following statements about human population in industrialized countries is
A) Average family size is relatively small.
B) The population has undergone the demographic transition.
C) Life history is r-selected.
D) The survivorship curve is Type I.
E) Age distribution is relatively uniform.
10) A recent study of ecological footprints (described in the text) concluded that
A) Earthʹs carrying capacity for humans is about 10 billion.
B) Earthʹs carrying capacity would increase if per capita meat consumption increased.
C) current demand by industrialized countries for resources is much smaller than the
ecological footprint of those countries.
D) the ecological footprint of the United States is large because per capita resource use is
E) it is not possible for technological improvements to increase Earthʹs carrying capacity
for humans.

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